Projects

The Center for Native Grasslands Management has undertaken a number of research initiatives focused on several key areas including biofuels, forage production, oak ecosystems, wildlife responses, and management of existing native grass stands.

Evaluation of Eastern Gamagrass and a Sorghum × Sudangrass for Summer Pasture

In the tall fescue belt, there is a well-known “summer slump” in forage production. This is because tall fescue, a cool-season grass, is less productive during summer. Warm-season grasses are needed that can fill this slump. One possibility is eastern gamagrass (EG), a highly productive warm-season perennial. In a four-year trial conducted at the Blount Unit of the East Tennessee Research and Education Center, bred yearling beef heifers grazed either EG or a commonly used summer annual, sorghum x sudangrass hybrid (SXS). During our trial, the grazing season for EG (112 days) lasted longer than the annual (65 days). The EG also allowed for higher stocking (1,591 vs. 1,239 lb per ac) and, as a result, produced more gain per acre (234 vs. 129 lb per acre) in the second and third years of the study. In the first year, rate of gain for EG was quite low (0.46 lb per day) leading to lower total gain per acre than for the SXS (108 vs. 149 lb per acre). 

Due to the need for annual establishment and the shorter grazing season, cost of gain for SXS ($1.67 per lb) was greater than that for EG ($0.62 per lb) except in the first year when the EG average daily gain was so low ($1.54 vs. $1.41 for SXS and EG, respectively).

Neither forage had any negative impact on pregnancy retention. This study, published in the Agronomy Journal, indicates that perennials have an advantage over annuals in terms of economics and length of grazing season. They also allow producers to avoid the risk of annual establishment and ensure a drought-tolerant forage is available throughout the summer grazing season.  

This study was supported by the University of Tennessee Institute of Agriculture and the USDA Cooperative State Research, Education, and Extension Service through Tennessee Hatch projects. 


Patch-burn Grazing Native Grasses in the Eastern US

Patch-burn grazing, also known as “pyric herbivory” is an approach for pasture management that mimics the ecological pattern once common in North American grasslands. Historically, bison followed the mosaic of burns across the prairies in search of the new, tender and nutritious forage that grew back following the fire.  In more recent years, researchers at Oklahoma State University began experimenting with burned patches within pastures. Using domestic cattle rather than bison, they burned 1/3 of each pasture annually. They found that the domestic cattle behaved much like bison – they spent most of their time grazing the most recently burned patch where the most nutritious forage was growing. As the patches were burned in successive years, the cattle also moved in response to the fire. Following this pattern, the grasses within patches burned in previous years received less grazing pressure and, as a result, were allowed to rest and restore their energy reserves.

We decided to try that here in the eastern US where pasture sizes are much smaller. At sites in Kentucky and Tennessee, we established 14 native grass pastures that averaged about 23 acres each. Seven of these pastures were assigned to a 3-paddock rotational system while the other seven were managed with the patch-burning. The burned pastures had 1/3 (about 8 ac) burned each spring, typically in early April. 

What did we learn? That patch-burn grazing can, in fact, work successfully at this scale. However, it was obvious that the difference in utilization of the current year’s burned patch was influenced a great deal by two factors: stocking rate and time between the burn and initiation of grazing.  Lighter stocking (increasing selectivity) and quicker stocking following the burn (greater nutritional quality of the burned forage) both increased the intensity of grazing of the burned patch.  Rates of gain, grazing days per acre, and overall gain per acre did not differ between the two approaches, rotational and patch-burning. 

This study was supported by a grant under the USDA Conservation Innovation Grant program administered by NRCS and was also supported by the Universities of Kentucky and Tennessee. 


Continuous Grazing of Mixed Native Warm-Season Grass in the Fescue Belt

Native grasses have often been characterized as being difficult to manage. The underlying concern is that they can be easily overgrazed and the stand quickly degraded. Therefore, rotational grazing has generally been recommended for managing native grass pastures. However, many stockmen in the Great Plains graze native grass rangelands all season long at set stocking rates. Because many producers may not want to follow strict rotational grazing guidelines, we wanted to explore how a minimal management approach could work for these grasses. Therefore, we initiated a study that examined two continuous grazing strategies, season-long, continuous grazing and a variation we called, “Heavy early.” The Heavy approach started the grazing season with 25% greater stocking than the simple continuous approach but, after late June, stocking was reduced to 25% below that of the simple continuous. This strategy was designed to more closely match the growth curves of native grasses, capture the spring flush and limit stress on the stand later in the summer.

After three summers we found that both approaches produced similar overall rates of gain for steers (2.06 lb per day), grazing days per acre (152 steer days), and total gain (317 lb per acre). More importantly though, we found that the sward vigor of the stands did not appear to be affected by the continuous grazing. At the conclusion of the study, tiller density (a reliable measure of stand vigor) was at 73 per sq foot indicating the stands had not been degraded. This despite an average of 105-day grazing season each summer and at least one severe drought (2016) during the study. Based on this work, it appears that managing native grass pastures can be quite simple and does not require precise rotational grazing or intensive management to be successful. 

This study was supported by the University of Tennessee Foundation through the gift of a generous, anonymous donor and by the University of Tennessee Institute of Agriculture. 


Impact of Perennial Warm-Season Grass Forages on Profitability of Beef Cattle Operations in the Fescue Belt

Although a number of studies have evaluated net returns from summer grazing and cost of gain under specific grazing strategies, there are still no data on the impact of an integrated warm-season/cool-season forage model on overall enterprise profitability. Ideally, such an evaluation should be based on empirical data, from a field experiment examining such an integrated production model. Unfortunately, data from such experiments are lacking. Therefore, to gain some insight into what the economic impact of incorporating native grasses into tall fescue forage systems might have, we developed an economic model. The model takes into account pasture costs, calving rates, calf weaning weights, animal nutritional needs, hay feeding costs, weather variability (i.e., drought frequency and severity), and market prices for calves and cull cows. All data were based on either University of Tennessee Extension budgets, existing market data, and historical weather records. Incorporating variability for input data, net present value calculations were run with 5,000 iterations. Models included three scenarios, a 100% fescue system and two integrated systems each with 30% warm-season grass, one with bermudgrass and one with switchgrass. For all three forage systems, we included a simulation with fall- and spring-calving herds. 

Based on this modeling exercise, cost of pastures was lowest for switchgrass and highest for bermudagrass, largely driven by input costs. Net present value, therefore, was always greatest for the switchgrass systems. For spring-calving herds, the bermudagrass system was also more profitable then the tall fescue only system. However, for fall-calving herds, the tall fescue only was more profitable than the bermudagrass system. While these results are based only on a model, and should be viewed cautiously, they do suggest that the incorporation of a low-cost, warm-season perennial such as switchgrass could make substantial contributions to enterprise profitability.   

This research was supported by the University of Tennessee Foundation through the gift of a generous, anonymous donor and by the University of Tennessee Institute of Agriculture. 

Using Nurse Crops During Establishment of Native Grass Forages

Native grasses can be challenging to establish. In addition, during the seedling year, use of stands for grazing or hay production is discouraged because of negative impacts on seedling development. Therefore, producers lose production for an entire year during the establishment process. One possible to solution to off-set this gap in production is to use a nurse crop during the establishment year. Planting a summer annual, but at a reduced density, could provide some competition to suppress weeds while also offering a forage crop.   

We began exploring this by conducting a small plot experiment at the Plant Science Unit of the East Tennessee Research and Education Center. We drilled browntop millet at a full rate and half rate into recently planted stands of switchgrass and big bluestem. During the summer, we harvested the millet based on two criteria – when the millet was ready for hay harvest and when the native grass seedlings were being overtopped by the millet. Unfortunately, our timing on these harvests were hampered by work schedules and weather. As a result, the millet only ended up adding to the overall level of competition for the native grass seedlings. 

Based on that first study, we have initiated a second study at the Greenacres Foundation in Ohio. In this second trial, we included an additional summer annual, a high diversity blend. In this project, we relied on grazing rather than mechanical removal of the summer annuals. During our first year, we wound up with a very similar problem, two much competition from the annuals as a result of our timing and intensity of grazing. As this project continues, we anticipate being able to adjust the grazing management to allow for a more favorable outcome. One thing appears to be obvious though. Trying to produce forage during the seedling year requires very good timing and implementation of canopy management treatments. There does not appear to be much room for error. This project will continue through 2021.   

These studies were supported by a USDA Agriculture and Food Initiative grant and a grant from the Greenacres Foundation. 


Overseeding Winter Annuals into Native Grasses

Although native grasses can produce large amounts of high quality forage during summer, many producers may want to extend the grazing seasons on these fields during the fall and/or spring. The obvious solution is to overseed the stands with a winter annual forage crop. After all, this is a widely used practice with other warm-season perennials such as bermudagrass. An initial evaluation was conducted using three winter annuals – cereal rye, wheat, and annual ryegrass. The winter annuals were subjected to one of three harvest timings – April 15, May 1, or May 15. Although we expected the latter maturing annuals and later harvest dates to have an increasingly negative effect on switchgrass stand vigor and yields, that was not the case. In fact, there was no loss in stand vigor (as measured by tiller density) or in subsequent switchgrass yields. 

Currently, we are wrapping up a grazing study in which existing switchgrass and big bluestem pastures have been overseeded with either cereal rye or a blend of rye, brassicas, and clovers. To date, it does not appear that the winter annuals have had any negative affect on the native grasses. However, we have also had some difficulty in getting strong stands of winter annuals due to late planting (first year of the experiment) and dry fall weather and/or exceptionally cold winters. This underscores the risk of relying on annual forage crops. This past fall, we got very good stands of annuals and some excellent grazing.  Analysis of results will continue through 2020. 

These projects were supported by grants from USDA Agriculture and Food Initiative and the USDA Cooperative State Research, Education, and Extension Service through Tennessee Hatch projects. 

Northern Bobwhite Survival and Habitat Use in Native Grass Pastures in the Fescue Belt

Northern bobwhite populations have experienced long-term, severe population declines. Past conservation efforts have focused disproportionately on habitat outside of production agriculture, on idled acres. A good example of this approach has been the Conservation Reserve Program (CRP), which takes land out of production. An alternative approach, one that could potentially impact many more acres, is a working lands model. Such a model focuses not on removing land from production, but rather on how to produce agricultural commodities and conservation benefit from the same land. One example of that is forage production using native grasses. Research has shown these grasses produce large volumes of loc-cost, high quality forage. What is less clear though, is how grassland wildlife species such as the northern bobwhite fare in when native grasses are subjected to grazing. Although studies in rangeland in the Great Plains suggest that such a combination is a “win-win”, very little data exists for eastern grasslands, such as those in the Tall Fescue Belt. 

Beginning in 2019, we initiated a study, the first of its kind east of the Mississippi, to evaluate bobwhite populations within grazed native grass systems. This radio telemetry study is examining habitat selection, nest survival, and adult survival throughout the year on the Blue Grass Army Depot in Kentucky. Although hampered by a small sample size, a result of a very small population (unfortunately, the norm in the Tall Fescue Belt in recent years), some interesting data is emerging. With no specific accommodation of bobwhite in the grazing regime, these birds are readily using, nesting in, and surviving in native grass pastures. By contrast, use of surrounding tall fescue pastures is very limited. The project is scheduled to continue through 2021. Populations of grassland passerines are also being monitored as a part of this study. 

Support for this research is being provided by a grant from the USDA, Natural Resources Conservation Service’s Working Lands for Wildlife Program, the Kentucky Department of Fish and Wildlife Resources, and the University of Tennessee Institute of Agriculture. 


Density and Nest Success of Grasshopper Sparrow and Field Sparrow on Continuously Grazed Native Warm-Season Grass Pastures

There is a lack of data on grassland bird response to grazing in native grass pastures. Given the potential benefits of a working lands conservation model for conservation of these at-risk species, understanding their response is critical. Therefore, we initiated a study on eight 20-acre native grass pastures in east Tennessee. All pastures were subjected to continuous, season long grazing (105-day duration) with one of two treatments: a standard stocking rate, and one that increased stocking by approximately 20% prior to late June, and then reduced that rate to 20% below the standard during late summer.

In addition, we monitored these same species on four reference tall fescue pastures of the same size and located in the same vicinity. Based on data collected over three summers, we learned that these species selected for the native grass pastures over tall fescue.

Although nest success and productivity were not different among the pasture treatments, the greater number of nests in the native grass pastures resulted greater overall productivity. Grasshopper produced 2 or 6 times and field sparrows 3 or 6 times more fledglings per acre compared with tall fescue pastures. The greater increases were associated with grazing management that maintained more optimum canopy heights – the standard stocking rate (about 900-1,00 lb per acre). Nest loss due to trampling was minimal. This study reinforces the idea that grazed native grasses are compatible with successful conservation of at-risk grassland birds. 

This project was supported by the University of Tennessee Foundation through the gift of a generous, anonymous donor, the Tennessee Wildlife Resources Agency, and the University of Tennessee Institute of Agriculture. 

Enhanced Biodiversity within Native Grass Pastures

Increased interest in soil health, wildlife conservation, and continuing declines in pollinator populations have all drawn attention to enhanced biodiversity within forage systems. Work on – and guidance for – improved plant diversity within native grass pastures has been limited. Information is needed to determine which native forbs and legumes can be successfully established within pasture systems, which will persist, not become pests, produce flowers that will benefit pollinators, and contribute to cattle forage production.  

To address these questions, we initiated a study at the University of Tennessee’s Research and Education Center at Greeneville. Two pastures, composed of lowland switchgrass and the second a big bluestem indiangrass blend, were overseeded with a 12-species blend of native forbs and legumes. The pastures were then sub-divided into 20 paddocks each and assigned to one of 5 grazing treatments. These treatments include no rest (grazed all season), no grazing (control) and another three with rest periods early, mid, and late season. 

We used this approach rather than adjusting stocking rates because experience has shown us that under light stocking rates, natives can easily become over-mature and thus limit forage quality and pasture productivity. Too, these treatments, in effect, mimic a rotational system in terms of the timing of rest intervals. Data continues to be collected on this study. Completion will be in fall of 2022.


Native Forb and Legume “Variety Trial”

A companion project to the pasture study mentioned above, this study is evaluating yield, persistence, flowering phenology, and forage nutritive qualities of 14 native species. Such baseline data will be helpful in understanding how to best integrate native forbs into forage production systems. Harvesting will begin in 2020 and is planned to continue for three summers. Species include coreopsis, prairie clover, Illinois bundle flower, partridge pea, sunflowers, black-eyed Susan, and cone flowers, among others. 

These two projects are being supported by the University of Tennessee Foundation through the gift of a generous, anonymous donor, a grant from the Greenacres Foundation, a grant from the USDA, Natural Resources Conservation Service, and the USDA Cooperative State Research, Education, and Extension Service through Tennessee Hatch projects.


Biodiversity, Soil Health, and Sustainability in Native Grasslands

Native grasses produce an abundance of forage but seem to do so with only modest fertility levels. What is the mechanism that allows for this unexpected productivity? Some possible explanations include symbiotic relationships with free-living soil microbes that fix nitrogen and mycorrhizae that supply phosphorous to the plants.  Another alternative is mineralization – the decomposition of the abundant below-ground biomass produced by the prodigious roots of these grasses. Related to this last point is the role of invertebrates in shredding the biomass, initiating the process by which the decomposition begins.   

To further explore these possibilities, a study was initiated to examine soil microbe communities, mycorrhizae root colonization, and invertebrate populations associated with native grass stands.

The native grasses have been subjected to varying levels of N and P fertilization (at higher levels, these may suppress soil microbes and mycorrhizae, respectively), hay harvest regimes, and grazing. Furthermore, the pastures have been treated with either a simple (cereal rye) or complex (rye, brassicas, clovers) overseeding or left as an unplanted control.  Analyses are underway as field work for this project concludes in 2020. 

This work has been supported by a grant from the USDA’s Agriculture and Food Initiative and by the University of Tennessee Institute of Agriculture.

Avian Ecology During Oak Savanna and Woodland Restoration

Woodlands and savannahs once were prevalent across much of the eastern United States. Like many other native grassland communities however, they have been almost completely eliminated and are now considered one of the most imperiled communities in the US. Associated wildlife species such as prairie warbler and red-headed woodpecker are also of high conservation concern. We conducted research over a 10-year period that included monitoring of bird populations. Analyses included occupancy, abundance, and for two species, prairie warbler and red-headed woodpecker, nest success. 

Results demonstrated that occupancy was strongly influenced by live basal area for many species. However, percent herbaceous ground cover appeared to be an even stronger indicator of occupancy.  Interestingly, across the range of basal area and herbaceous groundcover we evaluated, presence of early successional species increased markedly, while that for late-successional species declined only moderately. Thus, the bird community experienced overall increased the presence of at-risk species without substantial decrease in late successional species, many of which were not of high conservation priority. 

Indeed, analysis of abundance, when factored by Partners in Flight conservation priority scores indicated that stands undergoing restoration provided much greater conservation benefit than the closed-canopy stands. Nest success for red-headed woodpecker was quite high (>80% survival). Prairie warbler nest survival was variable with much lower rates the year of a prescribed burn and higher rates one year-post burning.  Notably, no nest of either species was found in our undisturbed control stands. Woodland and savannah restoration could be a valuable tool to not only restore a rare community, but could also contribute to improved conservation of many early successional bird species.  

This research was supported by grants from the USDI’s Joint Fire Science Program, North Carolina Wildlife Resources Commission, the National Wild Turkey Federation, Tennessee Wildlife Resources Agency, and the University of Tennessee Institute of Agriculture. 

Assessment of Wildlife Sustainability in Switchgrass Biofuel Feedstock Production

Prospects for large-scale biomass feedstock based on dedicated herbaceous crops such as switchgrass led to questions about how it would impact wildlife. Several years ago, the Department of Energy had forecast that as many as 35 million acres of such crops may be required to meet congressionally mandated energy targets. At that scale, implications of land conversion for breeding grassland birds, many of which are already considered at risk, could be substantial. We conducted a research project that monitored breeding bird occupancy and nest success over two summers at three locations. At each location (east Tennessee, northwest Pennsylvania, and western Kansas), we identified 10 production fields and 10 “matrix” fields. The matrix fields represented the prevailing land use for the site that would be most likely to be converted to energy crops. For Tennessee, these were tall fescue pastures, for Pennsylvania, cool-season pastures, and for Kansas, Conservation Reserve Program fields. We also collected data on use of the fields by pollinators, principally native bee species.

While analysis for these data sets is still being finalized, preliminary results for the bird occupancy suggest that responses varied by species. Red-winged blackbird, common yellowthroat, and dickcissel seemed to prefer the switchgrass, northern bobwhite, indigo bunting, and field sparrow appeared to use them equally, while meadowlark and grasshopper sparrow had greater occupancy in matrix fields. Results varied by location with landscape factors influencing occupancy. We continue to evaluate these data and are developing a spatially-explicit landscape model that can be used for evaluating biomass scenarios and their likely effect on many of these bird species. 

This research was supported by a USDA Agriculture and Food Initiative grant. 

Potential of Eastern Military Bases for Conservation of Grassland Bird

Building Partnerships for Regional Grassland Bird Conservation in the Southeastern US Department of Defense Legacy Resources Project 05-105

Native grasslands are one of the most endangered ecosystems in the eastern United States. The grassland bird species that are supported by these grasslands are also in dire need of conservation attention because of declining populations. Military installations provide important grassland habitat that serves a vital function for military training. As a result, eastern military installations have the opportunity to provide a critical role in conservation of grasslands and grassland bird communities.

Project Goals:

  • Document the potential of eastern military bases for grassland bird conservation
  • Document the status of grassland birds on one focal military installation (Fort Campbell)
  • Foster the development of regional partnerships for grassland bird conservation

Additional Resources

A map of the eastern US showing top military bases for grassland bird conservation.
Individual location numbers indicate more information on specific installations. Get more information on all military bases shown.

UT Theses

Giocomo, J. 2005. Ecology and population dynamics of grassland birds at Fort Campbell Military Reservation, KY. Ph.D. dissertation, University of Tennessee, Knoxville.

Hinnebusch, D. M. 2008. Nesting success and population densities of grassland birds in the breeding and wintering seasons in Tennessee and Kentucky. MS thesis, University of Tennessee, Knoxville. 148pp.

Moss, E. D. 2001. Distribution and reproductive success of native grassland birds in response to burning and field size at Fort Campbell Military Reservation: Special focus on Henslow’s and grasshopper sparrows. M.S. thesis, Univ. Tennessee, Knoxville. 161pp.

Project Reports

Giocomo, J. J. and D. A. Buehler. 2005. Conservation potential for grassland birds on eastern United States Department of Defense Installations. Final report delivered to Department of Defense Legacy Resources Program. 46 pp.

Minser, W. G., D. A. Buehler, and J. J. Giocomo. 2005. Management of grassland, savannah, and barrens habitat at Fort Campbell, Kentucky/Tennessee. Final report delivered to Department of Defense Legacy Resources Program. 45 pp.

Journal Articles

Giocomo, J. J., E. D. Moss, D. A. Buehler, and W. G. Minser. 2008. Nesting biology of grassland birds at Fort Campbell, Kentucky and Tennessee. Wilson Journal of Ornithology 120:111-119.

A map of the eastern US shows upland sandpiper habitat areas.

USFWS conservation status: Bird of Conservation concern/nation
Estimated world population: 350,000
Population management goal: Maintain current population
Breeding bird survey population trend (1966-2004): +0.7%
Threat to aircraft: Little threat due to small size, typical behavior (hiding in grassy areas) and tendency to remain on the ground. According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 44 Upland Sandpipers were identified and reported to the FAA out of ~1800 birds identified .

Management Recommendations:

  • Maintain large (>100 ha), contiguous tracts of prairie or grassland habitats.  Restore native grassland areas wherever possible.
  • Maintain grassland by burning, grazing, or haying every 2-3 years to prevent encroachment of woody vegetation.
  • Provide a mosaic of grassland habitat by disturbing about one third to half of the habitat each year.  Undisturbed patches are idea for nesting habitat while the disturbed areas are used for feeding and brood cover.
  • Allow some blocks of grassland to be left undisturbed each year to serve as nesting cover.
  • Avoid burning, mowing, or plowing during the nesting season  (May to early July).
  • Provide display perches, such as fence posts, rock piles, or tree stumps.
  • Burning should occur in the early spring (March – early April) before nesting starts or in the fall (September- October) especially for to control woody encroachment.
  • Encourage no-till or minimum-till practices instead of annual tillage practices, so that habitat is undisturbed during the nesting season in agricultural lease lands.

Dechant, J. A., M. F. Dinkins, D. H. Johnson, L. D. Igl, C. M. Goldade, B. D. Parkin, and B. R. Euliss.  2003.  Effects of management practices on grassland birds: Upland Sandpiper.  Northern Prairie Wildlife Research Center, Jamestown, ND.  Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/upsa/upsa.htm (Version 12DEC2003).

USFWS conservation status: Bird of Conservation concern/national
Partners in Flight watch list: No national status
Estimated world population: 1,300,000
Population management goal: Increase by 50%
Breeding bird survey population trend (1966-2004): -1.3%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 38 were identified and reported to the FAA out of ~1800 birds identified.

Management Recommendations:

  • Maintain a mosaic of grasslands and wetlands so that while some units are being treated to halt succession, other units are available. Treated units should be small (100-200 ha) to minimize the number of displaced nesting harriers. Untreated units should be large enough to meet the requirements of multiple female harriers during the nesting season (individual territories at least 150-260Ha).  In tallgrass areas, provide native and/or tame grasslands that have been recently (≤3 yr) idled.
  • Mow, burn, or graze fields every 2-5 yrs to maintain accumulations of residual vegetation preferred by Northern Harriers. Where natural vegetation has been destroyed by drainage, burning, tillage, overgrazing, or conversion to cropland, plant warm-season grasses and legumes.
  • Provide large areas (≥100 ha) of idle prairie and grassland with patches (< 0.25 Ha) of woody plants (i.e., Blackberry).
  • Avoid disturbing nesting areas during the breeding season, about April through July.
  • Minimize human disturbance near nests.  Do not use chemical pesticides in habitats used by harriers.

Dechant, J. A., M. L. Sondreal, D. H. Johnson, L. D. Igl, C. M. Goldade, M. P. Nenneman, and B. R. Euliss.  2003.  Effects of management practices on grassland birds: Northern Harrier.  Northern Prairie Wildlife Research Center, Jamestown, ND.  Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/noha/noha.htm (Version 12AUG2004).

A map of the eastern US shows short-eared owl habitat areas.

USFWS conservation status: Bird of Conservation concern/national
Partners in Flight watch list: Watch List Species
Estimated world population: 2,400,000
Population management goal: Increase by 100%
Breeding bird survey population trend (1966-2004): -4.8%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 36 were identified and reported to the FAA out of ~1800 birds identified at least to species group.

Management Recommendations:

  • Short-eared Owls are nomadic, they may be present only sporadically, but suitable habitat should be maintained.
  • Preserve and/or restore native grassland. Maintain a mosaic of grasslands and wetlands so that some management areas are available for nesting, while others are being treated to halt woody encroachment.
  • Burning, mowing, or grazing every 2-5 yr is recommended to maintain habitat for small mammal prey. Burn or mow to reduce grass height and maintain vegetation 30-40 cm tall.
  • To prevent mortality or injury from collisions with fences, remove unused fences. Increase visibility of fences by hanging pieces of ribbon or foil.

Dechant, J. A., M. L. Sondreal, D. H. Johnson, L. D. Igl, C. M. Goldade, M. P. Nenneman, and B. R. Euliss.  2003.  Effects of management practices on grassland birds: Short-eared Owl.  Northern Prairie Wildlife Research Center, Jamestown, ND.  Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/seow/seow.htm (Version 12DEC2003).

A map of the eastern US shows sedge wren habitat areas.

USFWS conservation status: Bird of Conservation concern/national
Partners in Flight watch list: No national status
Estimated world population: 6,500,000
Population management goal: Maintain current population
Breeding bird survey population trend (1966-2004): +1.9%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 0 were identified and reported to the FAA out of ~1800 birds identified at least to species group.

Management Recommendations:

  • Provide areas of tall, dense planted cover, such as big bluestem (Andropogon gerardii) or Indiangrass (Sorghastrum nutans). Suitable habitat also may be provided by areas dominated by reed canary grass (Phalaris arundinacea) and switchgrass (Panicum virgatum) if wet-prairie or sedge-meadow habitats are not available.
  • Avoid disturbance, such as mowing or herbicide spraying, during the breeding season. Due to a relatively long nesting season, delay mowing even longer than the date generally recommended for other passerines of 15 July.
  • Spray noxious weeds on a spot-by-spot basis, rather than on an entire-field basis.
  • Create a mosaic of burned and unburned areas to provide for both nesting and foraging needs.
  • Prevent encroachment by woody species in idle grassland by periodic disturbance (burning, mowing, or grazing) every 2-5 years, depending upon local conditions.
  • A rotational system of two or more management units may be most beneficial in providing distinct stands of grasses of various heights, but warm-season grasses should not be cut or grazed  <25 cm.

Dechant, J. A., M. L. Sondreal, D. H. Johnson, L. D. Igl, C. M. Goldade, B. D. Parkin, and B. R. Euliss.  2003.  Effects of management practices on grassland birds: Sedge Wren.  Northern Prairie Wildlife Research Center, Jamestown, ND.  Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/sewr/sewr.htm (Version 12DEC2003).

A map of the eastern US shows grasshopper sparrow habitat areas.

USFWS conservation status: Bird of Conservation concern/national (Endangered in FL)
Partners in Flight watch list: Stewardship Species
Estimated world population: 15,000,000
Population management goal: Increase by 100%
Breeding bird survey population trend (1966-2004): -3.8%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 8 were identified and reported to the FAA out of ~1800 birds identified at least to species group.

Management Recommendations:

  • Provide at least 30 ha of contiguous grassland (Grasshopper Sparrows are considered area sensitive species).  Grassland restoration areas should be >50 ha and preferably >100 ha in size.
  • Implement a rotational disturbance regime to maintain grassland habitat.  Burn every 2-5 years to control woody encroachment; shorter burn rotation in wetter areas (mesic conditions).
  • In some locations, mowing before 15 April can improve habitat, and may be preferable to prescribed burning.
  • To avoid destruction of nests, conduct management treatments before birds arrive in the spring (15 April) or after all young have fledged (1 September). Prescribed burn early spring (March to early April) or late fall (October and November).
  • Use a rotational burning program in which 20-30% of the site is burned each year. Management units should be at least 20-30 ha.  Never burn, mow, or otherwise disturb an entire area in one breeding season.
  • Modify intervals as needed to allow vegetation to recover between disturbances to provide suitable habitat while keeping succession in check.

Dechant, J. A., M. L. Sondreal, D. H. Johnson, L. D. Igl, C. M. Goldade, M. P. Nenneman, and B. R. Euliss.  2003.  Effects of management practices on grassland birds: Grasshopper Sparrow.  Northern Prairie Wildlife Research Center, Jamestown, ND.  Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/grsp/grsp.htm (Version 12AUG2004).

A map of the eastern US shows Henslow's sparrow habitat areas.

USFWS conservation status: Bird of Conservation concern/national
Partners in Flight watch list: Watch List Species
Estimated world population: 79,000
Population management goal: Increase by 100%
Breeding bird survey population trend (1966-2004): -8.6%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 0 were identified and reported to the FAA out of ~1800 birds identified at least to species group.

Management Recommendations:

  • Provide >30 ha of contiguous grassland.  If contiguous management units not available, provide a complex of smaller units located near enough to one another to facilitate colonization from adjacent territories in available habitat. Grassland restoration areas should be >50 ha and preferably >100 ha in size.
  • Never burn, mow, or otherwise disturb an entire area in one breeding season. Disturbance reduces available habitat for one or two growing seasons.
  • Implement a rotational disturbance regime to maintain grassland habitat. Burn every 2-5 years to control woody encroachment.
  • To avoid destruction of nests, conduct management treatments before birds arrive in the spring (15 April) or after all young have fledged (1 September). Prescribed burn early spring (March to early April) or late fall (October and November).
  • Use a rotational burning program in which 20-30% of the site is burned each year. Management units should be at least 20-30 ha.
  • Modify intervals as needed to allow vegetation to recover between disturbances to provide suitable habitat while keeping succession in check.

Herkert, J. R.  2003. Effects of management practices on grassland birds: Henslow’s Sparrow. Northern Prairie Wildlife Research Center, Jamestown, ND. Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/hesp/hesp.htm (Version 12DEC2003).

A map of the eastern US shows horned lark habitat areas.

USFWS conservation status: Common
Partners in Flight watch list: No national status
Estimated world population: 140,000,000
Population objective: Increase by 50%
Breeding bird survey population trend (1966-2004): -2.2%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 178 were identified and reported to the FAA out of ~1800 birds identified at least to species group.

Management Recommendations:

  • Create short, sparse vegetation by burning, and mowing.
  • Burn in the spring or late fall to reduce woody species and shrub growth
  • Maintain large (>100 Ha) grassland areas with a mosaic of grassland habitats and vegetation heights.
  • Road construction plans should consider the effects of roads on bird densities in rights-of-way and ≤500 m from rights-of-
    way

Dinkins, M. F., A. L. Zimmerman, J. A. Dechant, B. D. Parkin, D. H. Johnson, L. D. Igl, C. M. Goldade, and B. R. Euliss.  2003.  Effects of management practices on grassland birds: Horned Lark.  Northern Prairie Wildlife Research Center, Jamestown, ND.  Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/hola/hola.htm (Version 28MAY2004).

A map of the eastern US shows Le Conte's sparrow habitat areas.

USFWS conservation status: Bird of Conservation concern/national
Partners in Flight watch list: No national status
Notes: Mainly wintering populations in eastern U.S.
Estimated world population: 2,900,000
Population objective: Maintain or increase current populations
Breeding bird survey population trend (1966-2004): -0.5%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 0 were identified and reported to the FAA out of ~1800 birds identified at least to species group.

Management Recommendations:

  • Burn every 2-4 yr to control woody encroachment and reduce litter build-up.  Maintain tall grass throughout the winter and burn periodically to allow for foraging space in between grass clumps.   Avoid annual mowing or burning, which can destroy nests and reduce dense litter needed for nesting.
  • Timing and type of management must be adjusted according to regional differences and annual precipitation.
  • Do not leave habitat idle for so long as to allow over-accumulation of litter.

Dechant, J. A., M. L. Sondreal, D. H. Johnson, L. D. Igl, C. M. Goldade, A. L. Zimmerman, and B. R. Euliss.  2003.  Effects of management practices on grassland birds: Le Conte’s Sparrow.  Northern Prairie Wildlife Research Center, Jamestown, ND.  Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/lcsp/lcsp.htm (Version 12AUG2004).

A map of the eastern US shows vesper sparrow habitat areas.

USFWS conservation status: Common
Partners in Flight watch list: No national status
Estimated world population: 30,000,000
Population management goal: Increase by 50%
Breeding bird survey population trend (1966-2004): -1.1%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 5 were identified and reported to the FAA out of ~1800 birds identified at least to species group. 

Management Recommendations:

  • Burn in the spring before breeding territories are established or in late summer or early fall after birds have left the area to control woody encroachment. Do not burn all the habitat in any one year, burn up to 1/3 of the habitat each year.  Leave some woody vegetation in patches to serve as perches.
  • Burn or mow roadsides every 3-5 yr to maintain vegetation quality. To reduce nest losses, mow roadsides only in early spring or late summer. Retain fence lines along roadsides, especially in areas where forbs are sparse, to serve as perches.
  • Delay spraying pesticides and mowing until after July to avoid the peak nesting period.
  • Maintain fencerows adjacent to cropland  Near cropland, increase the proportion of fencerows that consist of both herbaceous and shrubby vegetation.
  • Use no-tillage or minimum-tillage methods to retain crop residue and waste grain on the surface of fields for birds to use in Agricultural lease areas.
  • Limit pesticide use in areas where Vesper Sparrows forage. Use only rapidly degrading chemicals of low toxicity at the lowest rates possible.

Dechant, J. A., M. F. Dinkins, D. H. Johnson, L. D. Igl, C. M. Goldade, and B. R. Euliss.  2003.  Effects of management practices on grassland birds: Vesper Sparrow.  Northern Prairie Wildlife Research Center, Jamestown, ND.  Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/vesp/vesp.htm (Version 28MAY2004).

A map of the eastern US shows savannah sparrow habitat areas.

USFWS conservation status: Common
Partners in Flight watch list: No national status
Notes:
Estimated world population:
 82,000,000
Population management goal: Increase by 50%
Breeding bird survey population trend (1966-2004): -0.8%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 44 were identified and reported to the FAA out of ~1800 birds identified at least to species group.

Management Recommendations:

  • Promote management or enhancement activities that increase the amount of contiguous grassland habitat and remove woody vegetation within and along the periphery of grassland fragments to discourage predators that may use woody vegetation as travel corridors and to enlarge the amount of interior grassland.
  • Aim grassland restorations at benefitting bird species most sensitive to habitat fragmentation; restorations should be ≥50 ha, preferably >100 ha. Where grassland restorations ≥30 ha are not possible, establish several small grasslands, 6-8 ha minimum size, within 0.4 km of each other, and using adjacent grassland habitats (e.g., pasture, hayland, waterway) as corridors among tracts.
  • Avoid disturbing (e.g., burning, mowing, moderate or heavy grazing) suitable habitat during the breeding season, approximately 1 May to 1 September. Treatments in nesting habitat should be delayed until after 1 September to prevent destruction of fledglings and renesting females.
  • Burn grasslands managed for breeding bird habitat in early spring (March to April) or late fall (October to November). In grasslands ≥50 ha, burn 25-30% annually; burn every 5-7 yr (Madden et al. 1999).
  • On airports not large enough to provide habitat for nesting birds (e.g., where all of the grassland available must be mowed to meet Federal Aviation Administration standards), mow and maintain grass short enough (<4 cm) to discourage nesting. This may cause birds to select alternative areas where nesting success would be higher.

Swanson, D. A. 2003. Effects of management practices on grassland birds: Savannah Sparrow. Northern Prairie Wildlife Research Center, Jamestown, ND.  Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/savs/savs.htm (Version 12AUG2004).

A map of the eastern US shows dickcissel habitat areas.

USFWS conservation status: Bird of Conservation concern/national
Partners in Flight watch list: Watch List Species
Estimated world population: 22,000,000
Population management goal: Increase by 50%
Breeding bird survey population trend (1966-2004): -0.9%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 1 were identified and reported to the FAA out of ~1800 birds identified at least to species group. 

Management Recommendations:

  • Provide >30 ha of contiguous grassland. If contiguous management units not available, provide a complex of smaller units located near enough to one another to facilitate colonization from adjacent territories in available habitat. Grassland restoration areas should be >50 ha and preferably >100 ha in size.
  • Never burn, mow, or otherwise disturb an entire area in one breeding season. Disturbance reduces available habitat for one or two growing seasons.
  • Implement a rotational disturbance regime to maintain grassland habitat. Burn every 2-5 years to control woody encroachment.
  • To avoid destruction of nests, conduct management treatments before birds arrive in the spring (15 April) or after all young have fledged (15 September). Prescribed burn early spring (March to early April) or late fall (October and November).
  • Use a rotational burning program in which 20-30% of the site is burned each year. Management units should be at least 20-30 ha.
  • Modify intervals as needed to allow vegetation to recover between disturbances to provide suitable habitat while keeping succession in check.
  • Shape, as well as area, of management units must be taken into consideration; perimeter-area ratio strongly influenced occurrence.
  • Adjust timing and type of management according to habitat. For example, xeric and mesic prairies may differ in rates of postburn litter accumulation, such that xeric prairies should be burned less frequently.
  • Prescribed burning in summer or fall or light disking of selected portions of individual fields can maintain mid-successional seral stages and increase coverage of tall forbs.
  • Do not mow later than mid-September in northern regions, because vegetation will not have time to recover before the winter or the following spring. Avoid mowing or eliminating forbs, brush, and hedgerows.
  • Provide areas of tall, dense planted cover. Allow retired agricultural fields to undergo some secondary succession. However, when succession begins to advance to the point of becoming unsuitable for breeding habitat, implement burning and/or grazing to control the growth of woody vegetation.

Dechant, J. A., M. L. Sondreal, D. H. Johnson, L. D. Igl, C. M. Goldade, A. L. Zimmerman, and B. R. Euliss.  2003.  Effects of management practices on grassland birds: Dickcissel.  Northern Prairie Wildlife Research Center, Jamestown, ND.  Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/dick/dick.htm (Version 12DEC2003).

A map of the eastern US shows bobolink habitat areas.

USFWS conservation status: Bird of Conservation concern/national
Partners in Flight watch list: Watch List Species
Estimated world population: 22,000,000
Population management goal: Increase by 50%
Breeding bird survey population trend (1966-2004): -0.9%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 1 were identified and reported to the FAA out of ~1800 birds identified at least to species group. 

Management Recommendations:

  • Provide >30 ha of contiguous grassland.  If contiguous management units not available, provide a complex of smaller units located near enough to one another to facilitate colonization from adjacent territories in available habitat. Grassland restoration areas should be >50 ha and preferably >100 ha in size.
  • Never burn, mow, or otherwise disturb an entire area in one breeding season. Disturbance reduces available habitat for one or two growing seasons.
  • Implement a rotational disturbance regime to maintain grassland habitat. Burn every 2-5 years to control woody encroachment.
  • To avoid destruction of nests, conduct management treatments before birds arrive in the spring (15 April) or after all young have fledged (15 September). Prescribed burn early spring (March to early April) or late fall (October and November).
  • Use a rotational burning program in which 20-30% of the site is burned each year. Management units should be at least 20-30 ha.
  • Modify intervals as needed to allow vegetation to recover between disturbances to provide suitable habitat while keeping succession in check.
  • Shape, as well as area, of management units must be taken into consideration; perimeter-area ratio strongly influenced occurrence.
  • Adjust timing and type of management according to habitat. For example, xeric and mesic prairies may differ in rates of postburn litter accumulation, such that xeric prairies should be burned less frequently.
  • Prescribed burning in summer or fall or light disking of selected portions of individual fields can maintain mid-successional seral stages and increase coverage of tall forbs.
  • Do not mow later than mid-September in northern regions, because vegetation will not have time to recover before the winter or the following spring. Avoid mowing or eliminating forbs, brush, and hedgerows.
  • Provide areas of tall, dense planted cover. Allow retired agricultural fields to undergo some secondary succession. However, when succession begins to advance to the point of becoming unsuitable for breeding habitat, implement burning and/or grazing to control the growth of woody vegetation.

Dechant, J. A., M. L. Sondreal, D. H. Johnson, L. D. Igl, C. M. Goldade, A. L. Zimmerman, and B. R. Euliss.  2003.  Effects of management practices on grassland birds: Dickcissel.  Northern Prairie Wildlife Research Center, Jamestown, ND.  Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/dick/dick.htm (Version 12DEC2003).

A map of the eastern US shows eastern meadowlark habitat areas.

USFWS conservation status: Common
Partners in Flight watch list: No national status (11)
Population objective: Maintain current population
Notes:
Estimated world population:
 10,000,000
Population management goal: Increase by 100%
Breeding bird survey population trend (1966-2004): -2.9%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 268 were identified and reported to the FAA out of 18,378 birds identified at least to species group. 

Management Recommendations:

  • Promote greater forb density and diversity in managed grasslands to improve overall habitat quality and provide food sources such as insects.  This may be accomplished by interseeding forb species in grassland plantings.
  • Limit the encroachment of woody vegetation. Remove woody vegetation within and along the periphery of grassland fragments to discourage predators that may use woody vegetation as travel corridors and to enlarge the amount of interior grassland.
  • Maintain a complex of burned and unburned habitats to provide a variety of grassland habitat types. Conduct prescribed burns in late spring on warm-season grasses to eliminate or reduce competition by cool-season grasses and weeds.
  • Burn prairie patches >80 ha on a rotation schedule, with 20-30% of area treated annually. Small, isolated prairie patches should not have more than 50-60% of the total area burned at a time. Work to create a mosaic of burned and unburned areas.
  • Burn every 2-5 yr and mow only at intervals of >3 yr.
  • Cool burns are optimal because some bunchgrasses and forbs will remain after the burn.
  • Provide periodic disturbances such as haying or grazing to increase floristic and structural diversity of seeded-native. Optimal mowing frequency may be every 3-5 yr in late summer, involving some kind of raking to reduce the litter layer. Delay burning and mowing within the breeding season to enhance suitable nesting habitat or to prevent nest destruction. If management is required to control weeds in Kansas, use spot mowing and spot spraying after 15 July to reduce nest destruction.
  • If an area needs to be mowed throughout the breeding season, lower the cutter height and mow frequently to discourage birds from nesting in the area and creating a population sink.

Hull, S. D. 2003. Effects of management practices on grassland birds: Eastern Meadowlark. Northern Prairie Wildlife Research Center, Jamestown, ND.  Northern Prairie Wildlife Research Center Online.  http://www.npwrc.usgs.gov/resource/literatr/grasbird/eame/eame.htm (Version 12DEC2003).

A map of the eastern US shows lapland longspur habitat areas.

USFWS conservation status: Common
Partners in Flight watch list: Stewardship species
Estimated world population:
 150,000,000
Population management goal: Maintain or increase current population
Christmas Bird Count (CBC) population trend (1966-2004): Stable

Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 6 were identified and reported to the FAA out of ~1800 birds identified at least to species group.

Habitat: human dominated agricultural landscapes.  Habitat currently extensive in NA and do not appear to be threatened.

Lapland Longspurs do not breed in the U.S., but tend to concentrate along waterways and other open ground away from forest habitats.  Winter habitats include grain stubble, grassy areas, and prairie remnants with little or no snow cover.  Heavy snow cover prevents access to seeds and will cause flocks to move to areas with less snow.

Lapland Longspurs are susceptible to mass kills during migration at night during winter storms.  Mass kills of over a 1,000 birds are associated with falling snow, foggy conditions, and isolated groups of lights (< 10 lights in a rural area) that attract the birds and cause them to hit man made-structures.   Lighting recommendations include using strobe lights instead of continuous lights on towers and down-shielding security lighting.  Major kills seem to be more associated with security lighting than tower kills.

Heavy pesticide use on crop seeds to reduce insect populations in agricultural fields should be avoided in agricultural-lease lands, because Lapland Longspurs use these fields during the winter and migration.  Mass kills have been reported for some kinds of pesticides.  These birds are especially susceptible during the spring migration period, because application of pesticides often happens during this same period when the birds are feeding on the seeds in the fields.

Hussell, D. J. T., and R. Montgomerie. 2002. Lapland Longspur (Calcarius lapponicus). In The Birds of North America, No. 656 (A. Poole and F. Gill, eds.). The Birds of North America, Inc. Philadelphia, PA.

A map of the eastern US shows snow bunting habitat areas.

USFWS conservation status: Common
Partners in Flight watch list: Stewardship species
Estimated world population:
 19,000,000
Population objective: Maintain current population
Population management goal: Maintain current population
Christmas Bird Count (CBC) population trend (1966-2004): Stable
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 86 were identified and reported to the FAA out of ~1800 birds identified at least to species group.

Snow Buntings forage mostly in agricultural fields during the winter, and heavy pesticide use may cause potential problems for survival.

Habitat: human dominated agricultural landscapes.  Habitat currently extensive in NA and do not appear to be threatened.Lyon, B. and R. Montgomerie. 1995. Snow Bunting and  McCay’s Bunting (Plectrophenax nivalis and Plectrophenax hyperboreus)In The Birds of North America, No. 198-199 (A. Poole and F. Gill, eds.).  The Birds of North America, Inc. Philadelphia, PA.

A map of the eastern US shows Bachman's sparrow habitat areas.

USFWS conservation status: Bird of Conservation concern/national
Partners in Flight watch list: Watch List Species (17)
Notes:
Estimated world population:
 250,000
Population objective: Increase by 100%
Breeding bird survey population trend (1966-2004): -2.0%
Threat to aircraft: According to FAA National Wildlife Strike Database from Jan 1990 through Sept 2005, 0 were identified and reported to the FAA out of ~1800 birds identified at least to species group.

A map of the eastern US shows breeding bird diversity by county.
Species richness of breeding grassland birds in the eastern US by county.

Range maps were compiled from Laughlin and Kibbe 1985, Illinois Department of Natural Resources 1986-1991, Adamus 1987, Andrle and Carroll 1988, Carolina Bird Club 1988-1995, Virginia Society of Ornithology 1989, Brewer et al. 1991, Peterjohn and Rice 1991, Brauning 1992, Enser 1992, Veit and Petersen 1993, Bevier 1994, Buckelew 1994, Foss 1994, Palmer-Ball 1996, Robbins 1996, Nicholson 1997, Castrale et al. 1998, Hess et al. 2000, Wiedenfeld and Swan 2000, Peterjohn 2001, Wisconsin Society for Ornithology 2002, Florida Fish and Wildlife Conservation Commission 2003, Turcotte and Watts 1999, Sauer et al. 2004.
A map of the eastern US shows grassland patches by state.
Proportion of existing large (>40 ha) grassland patches in the eastern US by county. Darker areas represent higher proportions. The red dots represent selected Department of Defense installations
A map of the eastern US shows wintering bird diversity by county.
Species richness of wintering grassland birds in the eastern US by county. 

Range maps were compiled from Christmas Bird Count summary range maps (Audubon Society 1959 – 1988, Root 1988).
A map of the eastern US shows open habitat spaces by state.
Proportion of open habitats (e.g., native grasslands, hay-fields, and other agricultural lands) in the eastern US by county. The open habitats represent potential areas for grassland restoration. Darker areas represent higher proportions. The red dots represent selected Department of Defense installations (see table 4 for installation names and identification numbers).